I thought it was all in there.
The problem with beta readers who already know a story is that they can understand inferences that aren't well explained.
Fortunately, not all my beta readers know this story in its various incarnations.
I'm getting feedback that tells me I didn't get everything out of my head and on to the paper. Of course I know why this character did that, but apparently I never shared the reasons behind the action. Which means I need to go back and add some backstory or build up. Dang.
When I go back through the scenes and realize what's been left out, it seems obvious. But until someone tells me what to look for, I don't see it. I know the story and the characters too well.
Thank goodness for my beta readers!